There was a special on a cable channel that escapes me now called, “Chimpanzee Family Fortunes.” While I found it fascinating, it spoke to a lot of what you describe in this piece. Maybe it is even the same group of chimps, because they also booted out their aggressive leader and eventually let him back in a few years later (in poor health and malnourished). When a female and her child were lost away from the group overnight in the forest, they were petrified of predators, because they cried all night.
What was clear from the documentary was that primates need groups to survive, but that groups were tricky to navigate. The “politics” of allies, enemies, and how chimps treat each other seemed particularly intense within the chimp “tribe.” I thought to myself if I were a chimp, I’d need an anxiety medication, because they were constantly in a state of maintaining group acceptance.